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Acestrorhynchus falcatus (Acfal_u3.jpg)
by Holm, E.

Size (cm):	13.5 SL
Locality:	Guyana,  Waini River
Sex/Stage:
Date:
Reference:
Source:
Remark:	1992

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Acestrorhynchus microlepis (Acmic_u0.jpg)
by Holm, E.


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Size (cm):	6.45 SL
Locality:	Guyana, Essequibo River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 64256

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Anablepsoides waimacui (Anwai_m0.jpg)
by Holm, E.

Size (cm):	4.8 SL
Locality:	Guyana, Potaro River
Sex/Stage:	male
Date:
Reference:
Source:
Remark:	ROM 61452

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Anchovia surinamensis (Ansur_u4.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Barama R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66179

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Anchoviella guianensis (Angui_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66180

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Boulengerella cuvieri (Boluc_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62515

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Boulengerella cuvieri (Boluc_u1.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62514

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Brachyplatystoma vaillantii (Brvai_u1.jpg)
by Holm, E.

Size (cm):	25.2 SL
Locality:	Guyana, Waini River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66400

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Cephalosilurus albomarginatus (Cealb_u1.jpg)
by Holm, E.

Size (cm):	8 SL
Locality:	Guyana, Potaro River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61482; dorsal and ventral view.

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Chasmocranus brevior (Chbre_u1.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Potaro R
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61489

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Colomesus asellus (Coase_u1.jpg)
by Holm, E.

Size (cm):	5 SL
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66467

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Colomesus psittacus (Copsi_u4.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Morika R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66474

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Crenicichla alta (Cralt_u6.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61917

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Crenicichla lugubris (Crlug_u1.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:	juvenile
Date:
Reference:
Source:
Remark:	ROM 61918

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Crenicichla lugubris (Crlug_u2.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61445

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Curimata cyprinoides (Cucyp_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Barama R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66601

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Curimata vittata (Cuvit_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:	juvenile
Date:
Reference:
Source:
Remark:	ROM 63292

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Curimatella immaculata (Cuimm_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66583

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Cyphocharax festivus (Cyfes_u2.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61332

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Cyphocharax helleri (Cyhel_u4.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Barama R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66596

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Doras carinatus (Docar_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62634

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Doras micropoeus (Domic_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62600

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Ossancora punctata (Dopun_u0.jpg)
by Holm, E.

Size (cm):	8 TL
Locality:	Peru, Pacallpa
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55611

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Eigenmannia macrops (Eimac_u0.jpg)
by Holm, E.

Size (cm):	13.1 SL
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62579

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Eigenmannia virescens (Eivir_u4.jpg)
by Holm, E.

Size (cm):	13.4 SL
Locality:	Guyana, Barama R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66512

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Electrophorus electricus (Elele_u5.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61622

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Eleotris amblyopsis (Elamb_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66484

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Geophagus brachybranchus (Gebra_j0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:	juvenile
Date:
Reference:
Source:
Remark:	ROM 61924

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Geophagus brachybranchus (Gebra_u4.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Mazaruni R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 68870

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Guianacara owroewefi (Guowr_j0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Potaro R.
Sex/Stage:	juvenile
Date:
Reference:
Source:
Remark:	ROM 61541

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Gymnorhamphichthys rondoni (Gyron_u1.jpg)
by Holm, E.

Size (cm):	15.5 TL
Locality:	Guyana, Essequibo River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62575

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Gymnotus anguillaris (Gyang_u2.jpg)
by Holm, E.

Size (cm):	5.5 TL
Locality:	Guyana, Tukeit Falls
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61476

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Gymnotus anguillaris (Gyang_u3.jpg)
by Holm, E.

Size (cm):	9.8 TL
Locality:	Guyana, Tukeit Falls,
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61335

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Gymnotus carapo (Gycar_u2.jpg)
by Holm, E.

Size (cm):
Locality:	Peru
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55533

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Hassar orestis (Haore_u1.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62599

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Chasmocranus longior (Chlon_u3.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61490

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Leptodoras linnelli (Lelin_u5.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62632

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Megalonema platycephalum (Mepla_u2.jpg)
by Holm, E.

Size (cm):	3.2 SL
Locality:	Guyana, Mazaruni R
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 69508

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Microglanis poecilus (Mipoe_u1.jpg)
by Holm, E.

Size (cm):	2.4 SL
Locality:	Guyana, Essequibo River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 60738

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Microphis brachyurus (Mibra_u4.jpg)
by Holm, E.

Size (cm):	14.2 SL
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 65539

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Moenkhausia browni (Mobro_u0.jpg)
by Holm, E.

Size (cm):	4.1 SL
Locality:	Guyana, Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61438

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Moenkhausia chrysargyrea (Mochr_u2.jpg)
by Holm, E.

Size (cm):	4.9 SL
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 64274

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Moenkhausia cotinho (Mocot_u0.jpg)
by Holm, E.

Size (cm):	4.6 SL
Locality:	Guyana,  Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61378

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Moenkhausia lepidura (Molep_u0.jpg)
by Holm, E.

Size (cm):	4.9 SL
Locality:	Guyana, Essequibo R.,
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 64676

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Moenkhausia megalops (Momeg_u0.jpg)
by Holm, E.

Size (cm):	5.3 SL
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 64276

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Moenkhausia simulata (Mosim_u0.jpg)
by Holm, E.

Size (cm):	6.8 SL
Locality:	Peru, Panguana Camp
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55738

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Myleus pacu (Mypac_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Myoglanis potaroensis (Mypot_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61479

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pachypops fourcroi (Pafou_u1.jpg)
by Holm, E.

Size (cm):	17.9 SL
Locality:	Guyana, Barama River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66283

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Parapristella aubynei (Paaub_u0.jpg)
by Holm, E.

Size (cm):	3.3 SL
Locality:	Guyana, Moruka R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 67877

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Parodon buckleyi (Pabuc_u8.jpg)
by Holm, E.

Size (cm):	12.6 SL
Locality:	Peru, Panguana
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55930

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Phenacogaster megalostictus (Phmeg_u2.jpg)
by Holm, E.

Size (cm):	4.2 SL
Locality:	Guyana, Potaro R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61442

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Piabucus dentatus (Piden_u1.jpg)
by Holm, E.

Size (cm):	6 SL
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodella cristata (Picri_u0.jpg)
by Holm, E.

Size (cm):	20.1 SL
Locality:	Guyana, Essequibo River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62337

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodella elongata (Pielo_u0.jpg)
by Holm, E.

Size (cm):	6.9 SL
Locality:	Peru, Tumbes River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66763

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodella gracilis (Pigra_u3.jpg)
by Holm, E.

Size (cm):	14.3 SL
Locality:	Peru, Llullapichis River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55793

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodella macturki (Pimac_u7.jpg)
by Holm, E.

Size (cm):	5.1 SL
Locality:	Guyana, Waini River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66455

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodus albofasciatus (Pialb_u2.jpg)
by Holm, E.

Size (cm):	15.6 SL
Locality:	Guyana, Essequibo River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62384

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodus blochii (Piblo_u2.jpg)
by Holm, E.

Size (cm):	18 SL
Locality:	Guyana, Waini River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66395

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Pimelodus ornatus (Piorn_u2.jpg)
by Holm, E.

Size (cm):	20.7 SL
Locality:	Guyana, Potaro River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61334

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Plagioscion squamosissimus (Plsqu_u3.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Barama River
Sex/Stage:
Date:
Reference:
Source:
Remark:	H92-064

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Polydactylus virginicus (Povir_u7.jpg)
by Holm, E.

Size (cm):	7.9 SL
Locality:	Guyana, Pomeroon R
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66489

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Poptella compressa (Pocom_u4.jpg)
by Holm, E.

Size (cm):	5.2 SL
Locality:	Guyana, Waini R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 67364

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Prochilodus nigricans (Prnig_u1.jpg)
by Holm, E.

Size (cm):	14 SL
Locality:	Peru
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55601

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Prochilodus rubrotaeniatus (Prrub_u4.jpg)
by Holm, E.

Size (cm):	20.5 SL
Locality:	Guyana, Essequibo River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 63605

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Psectrogaster essequibensis (Psess_j0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:	juvenile
Date:
Reference:
Source:
Remark:	ROM 63523

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Cephalosilurus albomarginatus (Cealb_u0.jpg)
by Holm, E.

Size (cm):	8 SL
Locality:	Guyana, Potaro River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61482

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Rhamdia xetequepeque (Rhxet_u0.jpg)
by Holm, E.

Size (cm):	7.2 SL
Locality:	Peru, Jequetepeque River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 70112, holotype.

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Rhynchodoras xingui (Rhxin_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 62601

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Laimosemion agilae (Riagi_u1.jpg)
by Holm, E.

Size (cm):	2.6 SL
Locality:	Guyana, Waini River
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66492

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Anablepsoides waimacui (Riwai_f0.jpg)
by Holm, E.

Size (cm):	4 SL
Locality:	Guyana, Potaro R.
Sex/Stage:	female
Date:
Reference:
Source:
Remark:	ROM 61452

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Apionichthys finis (Sofin_u0.jpg)
by Holm, E.

Size (cm):	5.9 SL
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61852.

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Steindachnerina guentheri (Stgue_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Barama R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 66597

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Steindachnerina guentheri (Stgue_j0.jpg)
by Holm, E.

Size (cm):
Locality:	Guyana, Waini R.
Sex/Stage:	juvenile
Date:
Reference:
Source:
Remark:	ROM 66589

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Synbranchus marmoratus (Symar_u3.jpg)
by Holm, E.

Size (cm):
Locality:	Peru, Panguana
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 55553

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Trichomycterus punctulatus (Trpun_u0.jpg)
by Holm, E.

Size (cm):
Locality:	Peru, Jequetepeque R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 51725

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.

Vandellia beccarii (Vabec_u0.jpg)
by Holm, E.

Size (cm):	4.4 SL
Locality:	Guyana, Essequibo R.
Sex/Stage:
Date:
Reference:
Source:
Remark:	ROM 61068

© ALL RIGHTS RESERVED. Image may not be used for any purpose without permission from the copyright holder.