Characterized by having escal morphology: escal bulb with small rounded, anteriorly to anteroventrally directed papilla on anterior midline, continuous internally with slightly larger posteriorly to posterodorsally directed papilla on dorsal midline; compressed posterior appendage sigmoidal in shape and usually twice as long as escal bulb; darkly pigmented swollen basal part of posterior appendage in specimens 84mm and larger; presence of filaments on posterior appendage, pigmented in specimens 84 mm and larger; dorsal fin rays 5-8; anal fin rays 4-6; pectoral fin rays 17-22 (Ref. 86949).