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References using Epinephelus tukula
Ref No. Description Page
000171  Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter,1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p.
000559  Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino,1984 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). 130;pl.116
001623  Smith, J.L.B. and M.M. Smith,1963 The fishes of Seychelles. Rhodes University, Grahamstown. 215 p. 15
001830  Eschmeyer, W.N.,1990 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p.
002334  Randall, J.E., G.R. Allen and R.C. Steene,1990 Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. 115
002872  Bianchi, G.,1985 FAO species identification sheets for fishery purposes. Field guide to the commercial marine and brackish-water species of Pakistan. Prepared with the support of PAK/77/033 and FAO (FIRM) Regular Programme. Rome: FAO. 200 p.
003153  Heemstra, P.C. and J.E. Randall,1984 Serranidae. In W. Fischer and G. Bianchi (eds.). FAO species identification sheets for fishery purposes. Western Indian Ocean (fishing area 51). Rome, FAO. Vol. IV. 241 p.
003670  van der Elst, R.,1981 A guide to the common sea fishes of southern Africa. C. Struik, Cape Town. 367 p.
003678  Torres, F.S.B. Jr.,1991 Tabular data on marine fishes from Southern Africa, Part I. Length-weight relationships. Fishbyte 9(1):50-53.
003807  Smith, A. and P. Dalzell,1993 Fisheries resources and management investigations in Woleai Atoll, Yap State, Federated States of Micronesia. Inshore Fish. Res. Proj., Tech. Doc., South Pacific Commission. Noumea, New Caledonia. 64 p.
004319  Heemstra, P.C. and J.E. Randall,1986 Serranidae. p. 509-537. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. 533
004787  Randall, J.E. and P.C. Heemstra,1991 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p.
004868  Shao, K.-T., S.-C. Shen, T.-S. Chiu and C.-S. Tzeng,1992 Distribution and database of fishes in Taiwan. p. 173-206. In C.-Y. Peng (ed.) Collections of research studies on 'Survey of Taiwan biological resources and information management'. Institute of Botany, Academia Sinica. Vol. 2. (in Chinese).
005213  Fischer, W., I. Sousa, C. Silva, A. de Freitas, J.M. Poutiers, W. Schneider, T.C. Borges, J.P. Feral and A. Massinga,1990 Fichas FAO de identificaçao de espécies para actividades de pesca. Guia de campo das espécies comerciais marinhas e de águas salobras de Moçambique. Publicaçao preparada em collaboraçao com o Instituto de Investigaçao Pesquiera de Moçambique, com financiamento do Projecto PNUD/FAO MOZ/86/030 e de NORAD. Roma, FAO. 1990. 424 p. 207
005222  Heemstra, P.C. and J.E. Randall,1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. 248;fig.429;pl.24d
005736  Hla Win, U.,1987 Checklist of fishes of Burma. Ministry of Livestock Breeding and Fisheries, Department of Fisheries, Burma.
009710  Lieske, E. and R. Myers,1994 Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Haper Collins Publishers, 400 p. 26.2
011441  Randall, J.E.,1995 Coastal fishes of Oman. University of Hawaii Press, Honolulu, Hawaii. 439 p. 136;fig.321
012484  van der Elst, R.,1993 A guide to the common sea fishes of southern Africa. (3rd Ed.). Struik Publishers, Cape Town. 398 p. 327
026367  Foale, S.,1998 What's in a name? An analysis of the West Nggela (Solomon Islands) fish taxonomy. SPC Traditional Marine Resource Management and Knowledge Information Bulletin No. 9:3-19. 9
027253  Lee, C. and Y. Sadovy,1998 A taste for live fish: Hong Kong's live reef fish market. Naga ICLARM Q. 21(2):38-42. 40
028618  Randall, J.E.,1997 Randall's tank photos. Collection of 10,000 large-format photos (slides) of dead fishes. Unpublished.
028620  Randall, J.E.,1997 Randall's underwater photos. Collection of almost 2,000 underwater photos (slides). Unpublished.
030573  Sommer, C., W. Schneider and J.-M. Poutiers,1996 FAO species identification field guide for fishery purposes. The living marine resources of Somalia. FAO, Rome. 376 p. 195
030874  Allen, G.R. and R.C. Steene,1988 Fishes of Christmas Island Indian Ocean. Christmas Island Natural History Association, Christmas Island, Indian Ocean, 6798, Australia. 197 p. 180
031411  Ganaden, S.R. and F. Lavapie-Gonzales,1999 Common and local names of marine fishes of the Philippines. Bureau of Fisheries and Aquatic Resources, Philippines. 385 p. 198
031517  Wu, H.L., K.-T. Shao and C.F. Lai (eds.),1999 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p.
033390  Fricke, R.,1999 Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez): an annotated checklist, with descriptions of new species. Koeltz Scientific Books, Koenigstein, Theses Zoologicae, Vol. 31:759 p. 200
034166  Lau, P.P.F. and L.W.H. Li,2000 Identification guide to fishes in the live seafood trade of the Asia-Pacific region. World Wide Fund for Nature, Hong Kong. 137 p. 58
036648  Randall, J.E. and K.K.P. Lim (eds.),2000 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. 610
036670  Anonymous,2000 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa.
037631  Zajonz, U., M. Khalaf and F. Krupp,2000 Coastal fish assemblages of the Socotra Archipelago. p.127-170. In Conservation and sustainable use of biodiversity of Socotra Archipelago: marine habitat, biodiversity and fisheries surveys and management. Progress Report of Phase III. Senckenberg Research Institute and Natural History Museum, Frankfurt. 159
038732  Anonymous,2001 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes.
039231  Heemstra, P.C. and J.E. Randall,1999 Serranidae. p. 2442-2547. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 4. Bony fishes part 2 (Mugilidae to Carangidae). Rome, FAO. 2069-2790 p.
041468  Kinch, J.,1999 Economics and environment in island Melanesia: a general overview of resource use and livelihoods on Brooker Island in the Calvados chain of the Louisiade Archipelago, Milne Bay Province, Papua New Guinea. A report prepared for Conservation International, Port Moresby, National Capital District, Papua New Guinea.
043538  Shao, K.-T., C.-P. Lin, L.-T. Ho and P.-L. Lin,1990 Study on the fish communities from northern and southern waters of Taiwan by analyzing the impingement data. J. Fish. Soc. Taiwan 17(2):73-90. 82
045255  Kapoor, D., R. Dayal and A.G. Ponniah,2002 Fish biodiversity of India. National Bureau of Fish Genetic Resources Lucknow, India.775 p. 399
047567  Allen, G.R. and M. Adrim,2003 Coral reef fishes of Indonesia. Zool. Stud. 42(1):1-72. 31
047843  Huang, Z.,2001 Marine species and their distribution in China's seas. p. 404-463. Vertebrata. Smithsonian Institution, Florida, USA. 598 p. 426
048497  Hardy, J.D. Jr. (comp.),2003 Coral reef fish species. NOAA\National Oceanographic Data Center. NODC Coral Reef Data and Information Management System. USA. 537 p. 138
048635  Kuiter, R.H. and T. Tonozuka,2001 Pictorial guide to Indonesian reef fishes. Part 1. Eels- Snappers, Muraenidae - Lutjanidae. Zoonetics, Australia. 1-302. 153;fig.
051015  Garpe, K.C. and M.C. Öhman,2003 Coral and fish distribution patterns in Mafia Island Marine Park, Tanzania: fish-habitat interactions. Hydrobiologia 498:191-211. 210
051471  Carl, H.,2003 Danish fish names. Zoological Museum of Copenhagen. Unpublished.
053568  Letourneur, Y., P. Chabanet, P. Durville, M. Taquet, E. Teissier, M. Parmentier, J.-C. Quéro and K. Pothin,2004 An updated checklist of the marine fish fauna of Reunion Island, south-western Indian Ocean. Cybium 28(3):199-216. 210
055073  Chen, C.-H.,2004 Checklist of the fishes of Penghu. FRI Special Publication No. 4. 175 p. 63
057683  Pereira, M.A.M.,2000 Preliminary checklist of reef-associated fishes of Mozambique. Maputo, Ministtry for the Coordination of Environmental Affairs (MICOA). 21 pp. 25
058078  Baissac, J. de B.,1990 SWIOP/WP/54 - Checklist of the marine fishes of Mauritius. RAF/87/008/WP/54/90 Regional Project for the Development & Management of Fisheries in the Southwest Indian Ocean.
058108  Chinese Academy of Fishery Sciences,2003 Chinese aquatic germplasm resources database.
058472  Heemstra, P.C., K. Hissmann, H. Fricke and M.J. Smale,2006 Fishes of the deep demersal habitat at Ngazidja (Grand Comoro) Island, Western Indian Ocean. South African J. Sci. 102(9/10):444-460. 452
075154  Hoese, D.F., D.J. Bray, J.R. Paxton and G.R. Allen,2006 Fishes. In Beasley, O.L. and A. Wells (eds.) Zoological Catalogue of Australia. Volume 35 Australia: ABRS & CSIRO Publishing, 2178 p. 1020
075549  FAO-FIES,2008 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from, 29 April 2008.
081519  Taquet, M. and A. Diringer,2007 Poissons de l'Océan Indien et de la Mer Rouge. Versailles (France): Éditions Quæ. 527 p. 162
083882  FAO-FIES,2010 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from, March 2010.
084159  Golani, D. and S. V. Bogorodsky,2010 The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa 2463:1-135. 24
087848  May, D.,2005 Folk taxonomy of reef fish and the value of participatory monitoring in Wakatobi National Park, southeast Sulawesi, Indonesia. SPC Traditional Marine Resource Management and Knowledge Information Bulletin 18: 18-35. 34
089707  Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra,2011 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group. 356 p. 253
089967  Taquet, M. and A. Diringer,1992 Mérous de l'Océan Indien. La Réunion: Azalées Editions. 103 p. 76; Photo, Painting
090062  FAO-FIES,2012 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from, March 2012.
090102  Allen, G.R. and M.V. Erdmann,2012 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. 280
095632  FAO-FIES,2014 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from, April 2014.
097629  Wickel, J., A. Jamon, M. Pinault, P. Durville and P. Chabanet,2014 Composition et structure des peuplements ichtyologiques marins de l’île de Mayotte (sud-ouest de l’océan Indien). Cybium 38(3):179-203. 193
101110  FAO-FIES,2015 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from, [accessed 13/04/2015].
103126  Koror Republic of Palau.,2012 The Rock Islands Southern Lagoon.,d.dGc [Accessed 08/07/2015].
115257  FAO-FIES,2017 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from (accessed 08/06/2017).
118272  FAO-FIES,2018 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps:// [accessed May, 2018]
118580  SCRFA, Science and Conservation of Fish Aggregations,2018 SCRFA database. Web online database. SCRFA. [2013-onwards].
118793  Allen, G.R. and M.V. Erdmann,2013 Coral reef fishes of Timor-Leste. p. 33-82. In G.R. Allen and M.V. Erdmann (eds). A rapid biological assessment of Timor-Leste. Conservation International. 61;AnnexI
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