| Citation | Niigata Prefecture, 2001, Red Data Book of Niigata.. Niigata Prefecture, Niigata. (http://www.pref.niigata.lg.jp/kankyokikaku/1214240790991.html; http://www.pref.niigata.lg.jp/HTML_Article/rdb12_gyorui.pdf) |
| Réf. | 95970 Niigata Prefecture, 2001 |
| Page | |
| Named Used as Valid | Gasterosteus aculeatus aculeatus |
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| Localité | |
| Distribution | Japan (Chubu: Niigata) |
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