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Raja clavata Linnaeus, 1758

Thornback ray
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Raja clavata   AquaMaps   Data sources: GBIF OBIS
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Image of Raja clavata (Thornback ray)
Raja clavata
Picture by Stergiou, K.I.


Spain country information

Common names: Arraizabala, Bastina rajada, Bramante
Occurrence: native
Salinity: marine
Abundance: | Ref:
Importance: | Ref:
Aquaculture: | Ref:
Regulations: | Ref:
Uses: no uses
Comments: Found in Galicia (Ref. 86578). Recorded from the Balearic Islands (Ref. 26178).
National Checklist:
Country Information: https://www.cia.gov/library/publications/resources/the-world-factbook/geos/sp.html
National Fisheries Authority:
Occurrences: Occurrences Point map
Main Ref: Muus, B. and P. Dahlström, 1978
National Database: ICTIMED

Common names from other countries

Classification / Names Tên thường gặp | Các synonym ( Các tên trùng) | Catalog of Fishes(Giống, Các loài) | ITIS | CoL | WoRMS | Cloffa

Lớp phụ Cá sụn (cá mập và cá đuối) (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates)
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335).
  More on author: Linnaeus.

Environment: milieu / climate zone / depth range / distribution range Sinh thái học

Biển gần đáy; Mức độ sâu 5 - 1020 m (Ref. 114953).   Subtropical; 70°N - 38°S, 25°W - 50°E

Sự phân bố Các nước | Các khu vực của FAO | Các hệ sinh thái | Những lần xuất hiện | Point map | Những chỉ dẫn | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Length at first maturity / Bộ gần gũi / Khối lượng (Trọng lượng) / Age

Maturity: Lm 76.6, range 47 - 87.5 cm
Max length : 105 cm TL con đực/không giới tính; (Ref. 53748); 139.0 cm TL (female); common length : 85.0 cm TL con đực/không giới tính; (Ref. 9987); Khối lượng cực đại được công bố: 18.0 kg (Ref. 35388); Tuổi cực đại được báo cáo: 15 các năm (Ref. 41305)

Short description Khóa để định loại | Hình thái học | Sinh trắc học

Các tia vây lưng cứng (tổng cộng) : 0; Tia cứng vây hậu môn: 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).

Sinh học     Tự điển (thí dụ epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019, 89020, 89022); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023). Maturity age 7-8 years (Ref. 114953). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311, 88171). Utilized fresh and frozen (Ref. 9987).

Life cycle and mating behavior Maturities | Sự tái sinh sản | Spawnings | Egg(s) | Fecundities | Ấu trùng

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).

Main reference Upload your references | Các tài liệu tham khảo | Người điều phối : McEachran, John | Người cộng tác

McEachran, J.D. and K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)

IUCN Red List Status (Ref. 130435)

  Near Threatened (NT) ; Date assessed: 03 October 2005

CITES (Ref. 128078)

Not Evaluated

CMS (Ref. 116361)

Not Evaluated

Threat to humans

  Harmless




Human uses

Các nghề cá: Tính thương mại; cá để chơi: đúng
FAO(Các nghề cá: Sản xuất; publication : search) | FishSource | Biển chung quanh ta

Thêm thông tin

Trophic ecology
Các loại thức ăn
Diet compositions
Food consumptions
Food rations
Các động vật ăn mồi
Ecology
Sinh thái học
Home ranges
Life cycle
Sự tái sinh sản
Maturities
Fecundities
Spawnings
Spawning aggregations
Egg(s)
Egg developments
Ấu trùng
Sự biến động ấu trùng
Anatomy
Gill areas
Não bộ
Otoliths
Physiology
Body compositions
Nutrients
Oxygen consumptions
Dạng bơi
Swimming speeds
Visual pigment(s)
Âm thanh của cá
Diseases / Parasites
Toxicities (LC50s)
Genetics
Di truyền
Electrophoreses
Heritabilities
Human related
Aquaculture systems
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Estimates based on models

Preferred temperature (Ref. 115969): 7.1 - 15.8, mean 10.7 (based on 474 cells).
Phylogenetic diversity index (Ref. 82804):  PD50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00200 (0.00175 - 0.00228), b=3.26 (3.23 - 3.29), in cm Total Length, based on LWR estimates for this species (Ref. 93245).
Mức dinh dưỡng (Ref. 69278):  3.8   ±0.2 se; based on diet studies.
Thích nghi nhanh (Ref. 120179):  thấp, thời gian nhân đôi của chủng quần tối thiểu là 4.5 - 14 năm (K=0.09-0.14; tm=10; tmax=23; Fec=150).
Prior r = 0.18, 95% CL = 0.12 - 0.26, Based on 4 full stock assessments.
Fishing Vulnerability (Ref. 59153):  High vulnerability (60 of 100).
Climate Vulnerability (Ref. 125649):  High vulnerability (60 of 100).
Price category (Ref. 80766):   Medium.