Africa: widespread in southern Africa from the Zambian Congo system (Ref. 52193) and the Cunene, Okavango (Ref. 7248, 11970, 52193, 120641), and Zambezi south to the Phongolo (Ref. 52193). Also present in the Cuanza (Ref. 11970, 120641) and Bengo in Angola (Ref. 11970). Absent from the lower Zambezi, Buzi, Pungwe and Save systems (Ref. 7248, 52193).