Classification List

Fish species numbers in the FishBase classification list


     The classification schema above genus level is limited in FishBase to classes, orders, families and subfamilies ranks (the latter not yet displayed). It has the advantage to keep it simple; it has the disadvantage to ignore some important clades in fish phylogeny like Chondrichthyes, Teleostei, Acanthomorpha, Percomorpha.

     In general, it follows Nelson's Fishes of the World (2006) with the noticeable exception of Elasmobranchii and Holocephali being elevated to class rank But at family level, FishBase may follow Eschmeyer's Catalog of Fishes that is updated more frequently to integrate new stable groupings.

     Number of taxa from Orders to Species are given for what they are, a useful tool for information and database management. The design of some analyses (so-called Biodiversity analyses) that uses these numbers must be carefully assessed before processing (see footnote for some critical points).

     Catalog of Fishes gives also number of species, including available names in the sense of the International Code of Zoological Nomenclature, which we don't have. Possible discrepancies between the species and genera numbers are explained in Bailly (2010). As Catalog of Fishes is updated ahead of FishBase, the numbers given there are in general more accurate.

     Wiley and Johnson published a new classification for Teleostei (2010). They propose to stabilize several hypotheses published recently or some (long) time ago by listing the known evidences (synapomorphies). FishBase will follow this new arrangement as soon as the ichthyological community endorses it, e.g., as published in a next version of Fishes of the World. In the meantime, to avoid confusion for non-specialists, we hold on the current one.

Note that the numbers of species given below does not take into account subspecies.

Total count of Genera: 5274
Total count of Species: 36535


Number of Valid
Kingdom Phylum
Class
#6
Order
#93
Family
#626
Orders Families Genera Species
Animalia
Chordata
Myxini 1 1 6 91
Myxiniformes 1 6 91
Myxinidae 6 91
Petromyzonti 1 3 11 50
Petromyzontiformes 3 11 50
Petromyzontidae 9 45
Geotriidae 1 2
Mordaciidae 1 3
Elasmobranchii 13 65 214 1294
Hexanchiformes 2 4 7
Hexanchidae 3 5
Chlamydoselachidae 1 2
Heterodontiformes 1 1 10
Heterodontidae 1 10
Orectolobiformes 7 13 45
Rhincodontidae 1 1
Parascylliidae 2 8
Brachaeluridae 1 2
Orectolobidae 3 12
Hemiscylliidae 2 17
Stegostomatidae 1 1
Ginglymostomatidae 3 4
Lamniformes 8 10 16
Mitsukurinidae 1 1
Carchariidae 1 2
Odontaspididae 1 2
Pseudocarchariidae 1 1
Megachasmidae 1 1
Alopiidae 1 3
Cetorhinidae 1 1
Lamnidae 3 5
Carcharhiniformes 12 53 309
Pentanchidae 11 111
Dichichthyidae 1 5
Scyliorhinidae 4 39
Atelomycteridae 3 13
Proscylliidae 3 7
Pseudotriakidae 3 5
Leptochariidae 1 1
Triakidae 9 47
Hemigaleidae 4 8
Carcharhinidae 11 59
Galeocerdonidae 1 1
Sphyrnidae 2 13
Squaliformes 6 22 144
Dalatiidae 7 10
Etmopteridae 4 54
Somniosidae 6 17
Oxynotidae 1 5
Centrophoridae 2 17
Squalidae 2 41
Echinorhiniformes 1 1 2
Echinorhinidae 1 2
Pristiophoriformes 1 2 10
Pristiophoridae 2 10
Squatiniformes 1 1 26
Squatinidae 1 26
Torpediniformes 5 16 73
Platyrhinidae 2 5
Narkidae 6 12
Narcinidae 5 31
Hypnidae 1 1
Torpedinidae 2 24
Rhinopristiformes 5 13 76
Trygonorrhinidae 3 8
Rhinobatidae 4 40
Rhinidae 3 11
Glaucostegidae 1 10
Pristidae 2 7
Rajiformes 4 40 314
Rajidae 18 165
Arhynchobatidae 14 114
Gurgesiellidae 3 19
Anacanthobatidae 5 16
Myliobatiformes 12 38 262
Zanobatidae 1 2
Hexatrygonidae 1 1
Dasyatidae 19 112
Potamotrygonidae 5 40
Urotrygonidae 2 20
Gymnuridae 1 16
Plesiobatidae 1 1
Urolophidae 3 28
Aetobatidae 1 5
Myliobatidae 2 20
Rhinopteridae 1 7
Mobulidae 1 10
Holocephali 1 3 6 58
Chimaeriformes 3 6 58
Callorhinchidae 1 3
Chimaeridae 2 46
Rhinochimaeridae 3 9
Cladistii 1 1 2 14
Polypteriformes 1 2 14
Polypteridae 2 14
Chondrostei 1 2 5 29
Acipenseriformes 2 5 29
Acipenseridae 3 27
Polyodontidae 2 2
Holostei 2 2 3 9
Lepisosteiformes 1 2 7
Lepisosteidae 2 7
Amiiformes 1 1 2
Amiidae 1 2
Teleostei 71 545 5023 34982
Hiodontiformes 1 1 2
Hiodontidae 1 2
Osteoglossiformes 6 31 262
Pantodontidae 1 1
Osteoglossidae 2 6
Arapaimidae 2 6
Notopteridae 4 10
Mormyridae 21 238
Gymnarchidae 1 1
Elopiformes 2 2 9
Elopidae 1 7
Megalopidae 1 2
Albuliformes 1 3 13
Albulidae 3 13
Notacanthiformes 2 6 30
Halosauridae 3 16
Notacanthidae 3 14
Anguilliformes 16 157 1040
Protanguillidae 1 1
Synaphobranchidae 12 51
Heterenchelyidae 2 8
Myrocongridae 1 6
Muraenidae 16 223
Chlopsidae 8 25
Colocongridae 1 11
Derichthyidae 2 3
Ophichthidae 63 361
Muraenesocidae 6 15
Nettastomatidae 6 47
Congridae 30 232
Moringuidae 2 18
Nemichthyidae 3 9
Serrivomeridae 2 10
Anguillidae 2 20
Saccopharyngiformes 5 5 28
Cyematidae 1 1
Monognathidae 1 15
Neocyematidae 1 1
Eurypharyngidae 1 1
Saccopharyngidae 1 10
Clupeiformes 10 85 441
Denticipitidae 1 1
Spratelloididae 2 8
Clupeidae 7 15
Dorosomatidae 30 115
Ehiravidae 11 31
Alosidae 4 32
Dussumieriidae 2 16
Engraulidae 18 183
Chirocentridae 1 2
Pristigasteridae 9 38
Alepocephaliformes 2 33 143
Alepocephalidae 20 103
Platytroctidae 13 40
Gonorynchiformes 4 7 41
Chanidae 1 1
Gonorynchidae 1 5
Kneriidae 4 34
Phractolaemidae 1 1
Cypriniformes 23 514 4832
Gyrinocheilidae 1 3
Catostomidae 15 82
Botiidae 8 62
Vaillantellidae 1 3
Cobitidae 20 238
Barbuccidae 1 3
Gastromyzontidae 20 152
Serpenticobitidae 1 3
Balitoridae 17 111
Ellopostomatidae 1 2
Nemacheilidae 45 807
Paedocyprididae 1 3
Psilorhynchidae 1 30
Cyprinidae 158 1787
Sundadanionidae 2 9
Danionidae 39 376
Leptobarbidae 1 5
Xenocyprididae 45 162
Tincidae 1 1
Acheilognathidae 7 82
Gobionidae 29 221
Tanichthyidae 1 6
Leuciscidae 99 684
Characiformes 28 291 2299
Citharinidae 3 8
Distichodontidae 16 110
Crenuchidae 12 106
Alestidae 19 120
Lepidarchidae 2 2
Hepsetidae 1 6
Tarumaniidae 1 1
Erythrinidae 3 20
Parodontidae 3 32
Cynodontidae 3 8
Serrasalmidae 17 104
Hemiodontidae 5 31
Anostomidae 16 165
Chilodontidae 2 8
Curimatidae 8 110
Prochilodontidae 3 21
Lebiasinidae 6 74
Ctenoluciidae 2 7
Chalceidae 1 7
Triportheidae 5 23
Gasteropelecidae 3 9
Bryconidae 4 49
Iguanodectidae 3 32
Acestrorhynchidae 7 26
Spintherobolidae 2 5
Stevardiidae 44 338
Characidae 38 207
Acestrorhamphidae 62 670
Gymnotiformes 5 34 262
Gymnotidae 2 45
Apteronotidae 15 98
Sternopygidae 6 51
Hypopomidae 6 38
Rhamphichthyidae 5 30
Siluriformes 41 516 4128
Nematogenyidae 1 1
Trichomycteridae 43 347
Callichthyidae 12 235
Loricariidae 115 1030
Scoloplacidae 1 6
Astroblepidae 1 81
Diplomystidae 2 7
Cetopsidae 5 43
Chacidae 1 4
Plotosidae 10 42
Ritidae 1 7
Ailiidae 5 17
Horabagridae 2 8
Bagridae 20 244
Auchenoglanididae 3 30
Akysidae 5 57
Amblycipitidae 4 42
Sisoridae 25 304
Pangasiidae 4 29
Siluridae 12 107
Kryptoglanidae 1 1
Aspredinidae 13 48
Auchenipteridae 26 127
Doradidae 31 97
Heptapteridae 23 226
Phreatobiidae 1 3
Pimelodidae 32 116
Pseudopimelodidae 7 53
Clariidae 16 118
Heteropneustidae 1 6
Ariidae 40 161
Anchariidae 2 6
Austroglanididae 1 3
Cranoglanididae 1 5
Ictaluridae 7 52
Lacantuniidae 1 1
Amphiliidae 13 102
Malapteruridae 2 21
Mochokidae 9 232
Claroteidae 8 62
Schilbeidae 9 47
Lepidogalaxiiformes 1 1 1
Lepidogalaxiidae 1 1
Argentiniformes 4 22 94
Argentinidae 2 28
Microstomatidae 3 21
Bathylagidae 8 23
Opisthoproctidae 9 22
Galaxiiformes 1 7 66
Galaxiidae 7 66
Esociformes 2 4 14
Esocidae 1 7
Umbridae 3 7
Salmoniformes 1 11 243
Salmonidae 11 243
Osmeriformes 4 17 42
Osmeridae 6 15
Plecoglossidae 1 1
Salangidae 7 20
Retropinnidae 3 6
Stomiiformes 4 52 447
Gonostomatidae 8 32
Sternoptychidae 10 79
Phosichthyidae 7 24
Stomiidae 27 312
Ateleopodiformes 1 4 14
Ateleopodidae 4 14
Aulopiformes 17 46 296
Aulopidae 4 16
Synodontidae 4 85
Pseudotrichonotidae 1 4
Paraulopidae 1 14
Chlorophthalmidae 2 21
Ipnopidae 5 31
Bathysauroididae 1 1
Scopelarchidae 5 18
Notosudidae 3 17
Giganturidae 1 2
Bathysauridae 1 2
Paralepididae 11 69
Anotopteridae 1 3
Evermannellidae 3 8
Omosudidae 1 1
Alepisauridae 1 2
Sudidae 1 2
Myctophiformes 2 37 257
Neoscopelidae 3 7
Myctophidae 34 250
Lampriformes 6 11 28
Lampridae 1 6
Veliferidae 2 2
Lophotidae 2 5
Radiicephalidae 1 2
Trachipteridae 3 10
Regalecidae 2 3
Percopsiformes 3 7 12
Percopsidae 1 2
Aphredoderidae 1 3
Amblyopsidae 5 7
Zeiformes 6 16 33
Cyttidae 1 3
Oreosomatidae 4 10
Parazenidae 3 4
Zeniontidae 3 7
Grammicolepididae 3 3
Zeidae 2 6
Stylephoriformes 1 1 1
Stylephoridae 1 1
Gadiformes 17 84 635
Bregmacerotidae 1 17
Phycidae 2 11
Gaidropsaridae 3 19
Lotidae 3 5
Gadidae 11 22
Ranicipitidae 1 1
Merlucciidae 2 17
Euclichthyidae 1 3
Muraenolepididae 2 9
Melanonidae 1 2
Trachyrincidae 4 9
Moridae 19 109
Macruronidae 1 4
Lyconidae 1 2
Bathygadidae 2 26
Steindachneriidae 1 1
Macrouridae 29 378
Polymixiiformes 1 1 12
Polymixiidae 1 12
Beryciformes 8 31 123
Berycidae 2 10
Melamphaidae 5 73
Stephanoberycidae 4 4
Hispidoberycidae 1 1
Gibberichthyidae 1 2
Rondeletiidae 1 2
Barbourisiidae 1 1
Cetomimidae 16 30
Trachichthyiformes 5 20 73
Monocentridae 2 5
Trachichthyidae 8 53
Anomalopidae 6 9
Diretmidae 3 4
Anoplogastridae 1 2
Holocentriformes 1 8 90
Holocentridae 8 90
Ophidiiformes 4 121 570
Ophidiidae 49 283
Carapidae 9 38
Bythitidae 38 135
Dinematichthyidae 25 114
Batrachoidiformes 1 23 83
Batrachoididae 23 83
Scombriformes 16 76 280
Amarsipidae 1 1
Nomeidae 3 16
Ariommatidae 1 7
Stromateidae 3 17
Pomatomidae 1 1
Centrolophidae 7 31
Icosteidae 1 1
Arripidae 1 4
Tetragonuridae 1 3
Chiasmodontidae 4 32
Scombridae 15 55
Caristiidae 4 19
Bramidae 7 20
Scombrolabracidae 1 1
Gempylidae 16 26
Trichiuridae 10 46
Syngnathiformes 5 67 339
Aulostomidae 1 3
Fistulariidae 1 4
Centriscidae 5 12
Solenostomidae 1 6
Syngnathidae 59 314
Dactylopteriformes 2 4 14
Pegasidae 2 7
Dactylopteridae 2 7
Callionymiformes 2 18 213
Callionymidae 16 198
Draconettidae 2 15
Mulliformes 1 6 104
Mullidae 6 104
Kurtiformes 2 41 369
Kurtidae 1 2
Apogonidae 40 367
Gobiiformes 13 324 2386
Trichonotidae 1 10
Rhyacichthyidae 2 3
Odontobutidae 7 25
Oxudercidae 10 42
Eleotridae 22 142
Butidae 11 53
Milyeringidae 2 6
Thalasseleotrididae 3 4
Gobiidae 244 1972
Kraemeriidae 2 9
Microdesmidae 13 92
Xenisthmidae 6 16
Schindleriidae 1 12
Synbranchiformes 4 17 131
Mastacembelidae 3 90
Chaudhuriidae 6 10
Indostomidae 1 3
Synbranchidae 7 28
Anabantiformes 8 26 281
Anabantidae 4 33
Helostomatidae 1 1
Osphronemidae 14 138
Aenigmachannidae 1 1
Channidae 2 56
Nandidae 1 7
Badidae 2 37
Pristolepididae 1 8
Carangaria/misc 8 17 109
Latidae 3 14
Centropomidae 1 13
Lactariidae 1 1
Sphyraenidae 1 29
Leptobramidae 1 2
Toxotidae 1 7
Menidae 1 1
Polynemidae 8 42
Pleuronectiformes 16 130 820
Psettodidae 1 3
Citharidae 4 6
Scophthalmidae 4 9
Cyclopsettidae 4 46
Bothidae 20 172
Paralichthyidae 10 65
Pleuronectidae 25 62
Paralichthodidae 1 1
Oncopteridae 1 1
Rhombosoleidae 8 20
Achiropsettidae 4 4
Achiridae 7 35
Samaridae 4 30
Poecilopsettidae 3 21
Soleidae 31 184
Cynoglossidae 3 161
Carangiformes 7 51 173
Nematistiidae 1 1
Carangidae 39 147
Echeneidae 3 8
Rachycentridae 1 3
Coryphaenidae 1 2
Istiophoridae 5 11
Xiphiidae 1 1
Cichliformes 2 247 1800
Cichlidae 246 1798
Pholidichthyidae 1 2
Atheriniformes 11 51 384
Atherinopsidae 13 114
Atherinidae 13 69
Atherionidae 1 3
Bedotiidae 2 16
Isonidae 1 5
Melanotaeniidae 7 114
Phallostethidae 4 24
Dentatherinidae 1 1
Pseudomugilidae 3 19
Telmatherinidae 5 18
Notocheiridae 1 1
Cyprinodontiformes 14 152 1442
Aplocheilidae 2 14
Nothobranchiidae 12 309
Rivulidae 48 469
Pantanodontidae 3 8
Fundulidae 3 44
Cyprinodontidae 9 103
Profundulidae 2 13
Goodeidae 18 51
Fluviphylacidae 1 8
Poeciliidae 28 277
Anablepidae 3 19
Aphaniidae 8 45
Valenciidae 1 3
Procatopodidae 14 79
Beloniformes 6 34 294
Adrianichthyidae 2 44
Scomberesocidae 2 5
Belonidae 10 49
Exocoetidae 7 71
Hemiramphidae 8 61
Zenarchopteridae 5 64
Mugiliformes 1 26 78
Mugilidae 26 78
Gobiesociformes 1 53 189
Gobiesocidae 53 189
Blenniiformes 6 153 946
Tripterygiidae 29 183
Labrisomidae 16 123
Clinidae 26 92
Chaenopsidae 14 98
Dactyloscopidae 9 48
Blenniidae 59 402
Ovalentaria/misc 8 96 849
Ambassidae 8 52
Embiotocidae 13 24
Grammatidae 2 18
Opistognathidae 4 112
Plesiopidae 12 51
Polycentridae 4 5
Pomacentridae 29 431
Pseudochromidae 24 156
Acanthuriformes 13 55 452
Lobotidae 3 15
Pomacanthidae 8 91
Drepaneidae 1 3
Ephippidae 8 15
Chaetodontidae 12 136
Leiognathidae 10 54
Scatophagidae 2 4
Antigoniidae 1 18
Siganidae 1 29
Caproidae 1 1
Luvaridae 1 1
Zanclidae 1 1
Acanthuridae 6 84
Lophiiformes 21 74 405
Lophiidae 4 34
Antennariidae 6 32
Histiophrynidae 7 15
Tathicarpidae 1 1
Tetrabrachiidae 2 2
Lophichthyidae 1 1
Rhycheridae 1 2
Brachionichthyidae 5 14
Chaunacidae 2 33
Ogcocephalidae 10 93
Caulophrynidae 2 5
Neoceratiidae 1 1
Melanocetidae 1 6
Himantolophidae 1 22
Diceratiidae 2 7
Oneirodidae 17 77
Centrophrynidae 1 1
Ceratiidae 2 4
Thaumatichthyidae 1 3
Gigantactinidae 2 25
Linophrynidae 5 27
Tetraodontiformes 10 105 450
Triodontidae 1 1
Triacanthodidae 11 23
Triacanthidae 4 7
Aracanidae 6 14
Ostraciidae 6 25
Diodontidae 7 18
Tetraodontidae 28 206
Monacanthidae 27 109
Balistidae 12 42
Molidae 3 5
Centrarchiformes 20 78 300
Percichthyidae 9 24
Terapontidae 16 59
Kuhliidae 1 12
Dichistiidae 1 2
Oplegnathidae 1 7
Kyphosidae 2 16
Girellidae 2 21
Scorpididae 4 11
Microcanthidae 4 6
Parascorpididae 1 1
Centrarchidae 8 41
Elassomatidae 1 7
Enoplosidae 1 1
Sinipercidae 2 14
Aplodactylidae 1 5
Cheilodactylidae 1 2
Chironemidae 1 6
Cirrhitidae 12 33
Latridae 9 30
Percalatidae 1 2
Acropomatiformes 20 58 305
Scombropidae 1 3
Ostracoberycidae 1 3
Acropomatidae 2 13
Symphysanodontidae 2 13
Epigonidae 7 50
Howellidae 3 9
Polyprionidae 1 2
Lateolabracidae 1 2
Glaucosomatidae 1 4
Pempheridae 2 80
Dinolestidae 1 1
Stereolepididae 1 2
Banjosidae 1 3
Pentacerotidae 7 13
Malakichthyidae 3 17
Champsodontidae 1 13
Hemerocoetidae 9 29
Creediidae 8 20
Synagropidae 4 18
Bathyclupeidae 2 10
Eupercaria/misc 19 274 1797
Moronidae 2 6
Dinopercidae 2 2
Centrogenyidae 1 1
Callanthiidae 2 18
Priacanthidae 4 22
Sillaginidae 5 38
Malacanthidae 2 16
Latilidae 3 30
Emmelichthyidae 3 19
Lutjanidae 21 138
Gerreidae 8 53
Haemulidae 21 138
Sparidae 38 163
Lethrinidae 5 43
Nemipteridae 5 77
Sciaenidae 68 300
Monodactylidae 1 4
Cepolidae 3 47
Labridae 80 682
Perciformes/Percoidei 3 14 254
Percidae 11 244
Trachinidae 2 9
Niphonidae 1 1
Perciformes/Zoarcoidei 14 108 418
Zoarcidae 57 300
Anarhichadidae 2 5
Neozoarcidae 2 10
Eulophiidae 3 5
Stichaeidae 18 45
Lumpenidae 8 11
Opisthocentridae 6 12
Pholidae 4 15
Ptilichthyidae 1 1
Zaproridae 1 1
Cryptacanthodidae 1 4
Cebidichthyidae 1 1
Scytalinidae 1 1
Bathymasteridae 3 7
Perciformes/Bembropoidei 1 2 23
Bembropidae 2 23
Perciformes/Percophoidei 1 1 1
Percophidae 1 1
Perciformes/Notothenioidei 8 46 161
Bovichtidae 3 11
Pseudaphritidae 1 2
Eleginopidae 1 1
Nototheniidae 13 56
Harpagiferidae 1 12
Artedidraconidae 5 36
Bathydraconidae 11 18
Channichthyidae 11 25
Perciformes/Uranoscopoidei 6 27 197
Trichodontidae 2 2
Ammodytidae 7 34
Uranoscopidae 7 57
Leptoscopidae 3 5
Pinguipedidae 7 98
Cheimarrichthyidae 1 1
Perciformes/Serranoidei 6 81 603
Serranidae 12 106
Anthiadidae 38 249
Epinephelidae 16 172
Grammistidae 8 33
Liopropomatidae 6 42
Caesioscorpididae 1 1
Perciformes/Scorpaenoidei 21 136 846
Sebastidae 7 133
Setarchidae 4 11
Neosebastidae 2 18
Scorpaenidae 28 241
Apistidae 3 4
Tetrarogidae 17 44
Synanceiidae 9 40
Eschmeyeridae 1 1
Perryenidae 1 1
Aploactinidae 17 50
Pataecidae 3 3
Gnathanacanthidae 1 1
Congiopodidae 1 6
Zanclorhynchidae 2 3
Bembridae 5 12
Plectrogeniidae 1 2
Hoplichthyidae 1 14
Platycephalidae 18 83
Triglidae 8 132
Peristediidae 6 46
Normanichthyidae 1 1
Perciformes/Gasterosteoidei 3 8 23
Hypoptychidae 2 2
Gasterosteidae 5 20
Aulorhynchidae 1 1
Perciformes/Cottoidei 10 158 882
Anoplopomatidae 2 2
Zaniolepididae 2 3
Hexagrammidae 3 9
Rhamphocottidae 3 4
Jordaniidae 3 3
Cottidae 72 294
Psychrolutidae 10 40
Agonidae 24 54
Cyclopteridae 7 31
Liparidae 32 442
Coelacanthi 1 1 1 2
Coelacanthiformes 1 1 2
Latimeriidae 1 2
Dipneusti 1 3 3 6
Ceratodontiformes 3 3 6
Neoceratodontidae 1 1
Lepidosirenidae 1 1
Protopteridae 1 4


Footnote: Minelli makes a definitive point about ranking as being an arbitrary and subjective decision: "The main difficulty resides in determining whether two clades in distant parts of the phylogenetic tree deserve [being] acknowledged [at] the same rank or not." (Minelli, 2009:11). "But all this amounts to counting fruit by adding apples and cherries." (Minelli, 2009:12). In other words, if counting orders, families, and genera in a narrow context (e.g., number of genera in subfamilies of the same family) may be locally sensible (within a phylogenetic tree / classification), counting families across classes or orders is most often irrelevant: for instance, why assigning clown-fishes only the subfamily rank (Pomacentridae: Amphiprioninae) while the Moorish idol has a family of its own (Zanclidae), and not a subfamily in Acanthuridae?

References:

Bailly, N., 2010. Why there may be discrepancies in the assessment of scientific names between the Catalog of Fishes and FishBase. Electronic document. [Version 2, May 6th, 2010]. //www.fishbase.org/Nomenclature/FBCofFNames.php

Eschmeyer, W.N. (ed.). Catalog of Fishes. Successive updated online versions http://research.calacademy.org/research/Ichthyology/catalog/fishcatmain.asp.

Eschmeyer, W.N. and J.D. Fong. Species of Fishes by family/subfamily. Successive updated online versions. http://research.calacademy.org/research/ichthyology/catalog/SpeciesByFamily.asp.

Minelli, A., 2009. Perspectives in animal phylogeny. Oxford University Press.

Nelson, J.S., 2006. Fishes of the World. 4th ed. Hoboken (New Jersey, USA): John Wiley & Sons. xix+601 p. [Ref. 58010]

Wiley, E.O. and G.D. Johnson, 2010. pp. 123-182. A teleost classification based on monophyletic groups. In: Origin and phylogenetic interrelationships of Teleosts. Honoring Gloria Arratia. Proceedings of the International Symposium at the ASIH Annual Meeting in St. Louis, Missouri, 2007. (Nelson et al., eds). München (Germany): Verlag Dr. Friedrich Pfeil. [Ref. 84927]

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