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<taxon>
	<dc:identifier>FB-2420</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2420</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Gasterosteoidei</dwc:Order>
	<dwc:Family>Gasterosteidae</dwc:Family>
	<dwc:Genus>Gasterosteus</dwc:Genus>
	<dwc:ScientificName>Gasterosteus aculeatus Linnaeus, 1758</dwc:ScientificName>
	<commonName xml:lang='no'>Stikling</commonName>
	<commonName xml:lang='no'>Trepigget stingsild</commonName>
	<commonName xml:lang='fa'>Sehkhareh</commonName>
	<commonName xml:lang='pl'>Ciernik</commonName>
	<commonName xml:lang='pt'>Esganagata</commonName>
	<commonName xml:lang='pt'>Esgana-gata</commonName>
	<commonName xml:lang='pt'>Espinhela</commonName>
	<commonName xml:lang='pt'>Peixe-espinho</commonName>
	<commonName xml:lang='sq'>Gjëmbaçi</commonName>
	<commonName xml:lang='sq'>Peshk koçkuni</commonName>
	<commonName xml:lang='bg'>Kedrinka</commonName>
	<commonName xml:lang='cs'>Koljueka triostna</commonName>
	<commonName xml:lang='cs'>Koljuška tríostná</commonName>
	<commonName xml:lang='cs'>Koljuška tříostná</commonName>
	<commonName xml:lang='cy'>Crothell dri phigyn</commonName>
	<commonName xml:lang='da'>Trepigget hundestejle</commonName>
	<commonName xml:lang='de'>Dreistacheliger Stichling</commonName>
	<commonName xml:lang='de'>Dreistachliger Stichling</commonName>
	<commonName xml:lang='de'>Großer Stichling</commonName>
	<commonName xml:lang='de'>Rotzbarsch</commonName>
	<commonName xml:lang='de'>Seestichling</commonName>
	<commonName xml:lang='de'>Stachel de butz</commonName>
	<commonName xml:lang='de'>Stachelbauch</commonName>
	<commonName xml:lang='de'>Stachele</commonName>
	<commonName xml:lang='de'>Stachelfisch</commonName>
	<commonName xml:lang='de'>Stechbüttel</commonName>
	<commonName xml:lang='de'>Stecherling</commonName>
	<commonName xml:lang='de'>Stechert</commonName>
	<commonName xml:lang='de'>Steckbedel</commonName>
	<commonName xml:lang='de'>Steckbüdel</commonName>
	<commonName xml:lang='de'>Steckelbars</commonName>
	<commonName xml:lang='de'>Steckelstange</commonName>
	<commonName xml:lang='de'>Steckker</commonName>
	<commonName xml:lang='de'>Steckling</commonName>
	<commonName xml:lang='de'>Stichling</commonName>
	<commonName xml:lang='en'>Banstickle</commonName>
	<commonName xml:lang='en'>Barnstickle</commonName>
	<commonName xml:lang='en'>Barnystickle</commonName>
	<commonName xml:lang='en'>Barnytickle</commonName>
	<commonName xml:lang='en'>Branchy</commonName>
	<commonName xml:lang='en'>Branstickle</commonName>
	<commonName xml:lang='en'>Burnstickle</commonName>
	<commonName xml:lang='en'>Common stickleback</commonName>
	<commonName xml:lang='en'>Cushy</commonName>
	<commonName xml:lang='en'>Doctor</commonName>
	<commonName xml:lang='en'>Eastern stickleback</commonName>
	<commonName xml:lang='en'>European stickleback</commonName>
	<commonName xml:lang='en'>Jacksharp</commonName>
	<commonName xml:lang='en'>New York stickleback</commonName>
	<commonName xml:lang='en'>Pinfish</commonName>
	<commonName xml:lang='en'>Prickley</commonName>
	<commonName xml:lang='en'>Prickly</commonName>
	<commonName xml:lang='en'>Prickly back</commonName>
	<commonName xml:lang='en'>Santa Ana stickleback</commonName>
	<commonName xml:lang='en'>Saw-finned stickleback</commonName>
	<commonName xml:lang='en'>Sparnicle</commonName>
	<commonName xml:lang='en'>Sparny</commonName>
	<commonName xml:lang='en'>Sparnytickle</commonName>
	<commonName xml:lang='en'>Spanicle</commonName>
	<commonName xml:lang='en'>Spannistickle</commonName>
	<commonName xml:lang='en'>Spanny</commonName>
	<commonName xml:lang='en'>Spannytickle</commonName>
	<commonName xml:lang='en'>Spantickle</commonName>
	<commonName xml:lang='en'>Spawn</commonName>
	<commonName xml:lang='en'>Spawnykettle</commonName>
	<commonName xml:lang='en'>Spawnytickle</commonName>
	<commonName xml:lang='en'>Stickleback</commonName>
	<commonName xml:lang='en'>Thorny back</commonName>
	<commonName xml:lang='en'>Thornback</commonName>
	<commonName xml:lang='en'>Threespine stickleback</commonName>
	<commonName xml:lang='en'>Threespined stickleback</commonName>
	<commonName xml:lang='en'>Three-spined stickleback</commonName>
	<commonName xml:lang='en'>Tiddler</commonName>
	<commonName xml:lang='en'>Twospine stickleback</commonName>
	<commonName xml:lang='en'>Unarmored threespine stickleback</commonName>
	<commonName xml:lang='et'>Harilik ogalik</commonName>
	<commonName xml:lang='et'>ogalik</commonName>
	<commonName xml:lang='et'>Paljasogalik</commonName>
	<commonName xml:lang='fo'>Beinmurtur</commonName>
	<commonName xml:lang='fo'>Eldmurtur</commonName>
	<commonName xml:lang='fo'>Eyrriðasild</commonName>
	<commonName xml:lang='fo'>Horngeil</commonName>
	<commonName xml:lang='fo'>Horngeir</commonName>
	<commonName xml:lang='fo'>Hvasssíl</commonName>
	<commonName xml:lang='fo'>Kombikk</commonName>
	<commonName xml:lang='fi'>Kolmipiikki</commonName>
	<commonName xml:lang='fr'>Arselet</commonName>
	<commonName xml:lang='fr'>Cordonnier</commonName>
	<commonName xml:lang='fr'>Crève-valet</commonName>
	<commonName xml:lang='fr'>Épinart</commonName>
	<commonName xml:lang='fr'>Épinglet</commonName>
	<commonName xml:lang='fr'>Épinoche</commonName>
	<commonName xml:lang='fr'>Epinoche à trois épines</commonName>
	<commonName xml:lang='fr'>Épinoche à trois épines</commonName>
	<commonName xml:lang='fr'>Estancelin</commonName>
	<commonName xml:lang='fr'>Estranglo cat</commonName>
	<commonName xml:lang='fr'>Spinaubé</commonName>
	<commonName xml:lang='fr'>Spinavaou</commonName>
	<commonName xml:lang='fr'>Stichling</commonName>
	<commonName xml:lang='hr'>Koljuška</commonName>
	<commonName xml:lang='hu'>Tükes pikós</commonName>
	<commonName xml:lang='is'>Hornsíli</commonName>
	<commonName xml:lang='iu'>Kakelashuk</commonName>
	<commonName xml:lang='iu'>Kakilaychok</commonName>
	<commonName xml:lang='iu'>Kakilisak</commonName>
	<commonName xml:lang='iu'>Kakilishek</commonName>
	<commonName xml:lang='iu'>Kakilusuk</commonName>
	<commonName xml:lang='iu'>Kakkilasak</commonName>
	<commonName xml:lang='iu'>Katilautik</commonName>
	<commonName xml:lang='it'>Spinarello</commonName>
	<commonName xml:lang='ja'>Hariyo</commonName>
	<commonName xml:lang='ja'>Itoyo</commonName>
	<commonName xml:lang='ja'>イトヨ</commonName>
	<commonName xml:lang='ko'>큰가시고기</commonName>
	<commonName xml:lang='lv'>trīsadatu stagars</commonName>
	<commonName xml:lang='lt'>trispyglė dyglė</commonName>
	<commonName xml:lang='ru'>Trekhiglaja koliushka</commonName>
	<commonName xml:lang='ru'>Trekhiglaya kolyushka</commonName>
	<commonName xml:lang='ru'>Колюшка трехиглая</commonName>
	<commonName xml:lang='ru'>трехиглая колюшка</commonName>
	<commonName xml:lang='sr'>Bodonja</commonName>
	<commonName xml:lang='sk'>Pichlavka</commonName>
	<commonName xml:lang='sk'>Pichlavka siná</commonName>
	<commonName xml:lang='sv'>Nakenspigg</commonName>
	<commonName xml:lang='sv'>Storspigg</commonName>
	<commonName xml:lang='tr'>Dikence baligi</commonName>
	<commonName xml:lang='tr'>Dikence balığı</commonName>
	<commonName xml:lang='uk'>Trigolkova koluchka</commonName>
	<commonName xml:lang='es'>Espinocho</commonName>
	<commonName xml:lang='es'>Espinosillo</commonName>
	<commonName xml:lang='es'>Espinoso</commonName>
	<commonName xml:lang=''>Agathero</commonName>
	<commonName xml:lang=''>Cukilrua'ak</commonName>
	<commonName xml:lang=''>Driedoornige stekelbaars</commonName>
	<commonName xml:lang=''>Espinós</commonName>
	<commonName xml:lang=''>Ghidrin</commonName>
	<commonName xml:lang=''>kakalisauraq</commonName>
	<commonName xml:lang=''>Snathaid mhara</commonName>
	<commonName xml:lang=''>Stekelbaars</commonName>
	<commonName xml:lang=''>Zet</commonName>
	<commonName xml:lang=''>Αγκαθερό</commonName>
	<commonName xml:lang=''>Σκυλόψαρο</commonName>
	<commonName xml:lang=''>Σταυρίδι</commonName>
	<commonName xml:lang=''>سه خاره</commonName>
	<commonName xml:lang=''>三刺魚</commonName>
	<commonName xml:lang=''>三刺鱼</commonName>
	<commonName xml:lang=''>威氏三刺魚</commonName>
	<commonName xml:lang=''>威氏三刺鱼</commonName>
	<commonName xml:lang=''>桑塔三刺魚</commonName>
	<commonName xml:lang=''>桑塔三刺鱼</commonName>
	<commonName xml:lang=''>阿拉斯加刺魚</commonName>
	<commonName xml:lang=''>阿拉斯加刺鱼</commonName>
	<synonym relationship='synonym'>Gasteracanthus cataphractus Pallas, 1814</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus aculeatus Linnaeus, 1758</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus algeriensis Sauvage, 1874</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus gymnurus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus leiurus Cuvier, 1829</synonym>
	<synonym relationship='ambiguous synonym'>Gasterosteus aculeatus messinicus Stephanidis, 1971</synonym>
	<synonym relationship='misapplied name'>Gasterosteus aculeatus microcephalus (non Girard, 1854)</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus santaeannae Regan, 1909</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus trachurus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus aculeatus williamsoni Girard, 1854</synonym>
	<synonym relationship='synonym'>Gasterosteus algeriensis Sauvage, 1874</synonym>
	<synonym relationship='synonym'>Gasterosteus argentatissimus Blanchard, 1866</synonym>
	<synonym relationship='synonym'>Gasterosteus argyropomus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus atkinsi Bean, 1879</synonym>
	<synonym relationship='synonym'>Gasterosteus atkinsii Bean, 1879</synonym>
	<synonym relationship='synonym'>Gasterosteus bailloni Blanchard, 1866</synonym>
	<synonym relationship='synonym'>Gasterosteus biaculeatus Mitchill, 1815</synonym>
	<synonym relationship='synonym'>Gasterosteus biarmatus Krynicki, 1840</synonym>
	<synonym relationship='synonym'>Gasterosteus bispinosus Walbaum, 1792</synonym>
	<synonym relationship='synonym'>Gasterosteus brachycentrus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus cataphractus (Pallas, 1814)</synonym>
	<synonym relationship='synonym'>Gasterosteus cuvieri Girard, 1850</synonym>
	<synonym relationship='synonym'>Gasterosteus Dekayi Ayres, 1855</synonym>
	<synonym relationship='synonym'>Gasterosteus dimidiatus Reinhardt, 1837</synonym>
	<synonym relationship='synonym'>Gasterosteus elegans Blanchard, 1866</synonym>
	<synonym relationship='synonym'>Gasterosteus gymnurus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus hologymnus Regan, 1909</synonym>
	<synonym relationship='synonym'>Gasterosteus inopinatus Girard, 1854</synonym>
	<synonym relationship='synonym'>Gasterosteus insculptus Richardson, 1855</synonym>
	<synonym relationship='synonym'>Gasterosteus intermedius Girard, 1856</synonym>
	<synonym relationship='misapplied name'>Gasterosteus islandicus (non Sauvage, 1874)</synonym>
	<synonym relationship='synonym'>Gasterosteus leiurus Cuvier, 1829</synonym>
	<synonym relationship='ambiguous synonym'>Gasterosteus loricatus Reinhardt, 1837</synonym>
	<synonym relationship='misapplied name'>Gasterosteus microcephalus (non Girard, 1854)</synonym>
	<synonym relationship='synonym'>Gasterosteus nemausensis Crespon, 1844</synonym>
	<synonym relationship='synonym'>Gasterosteus neoboracensis DeKay, 1842</synonym>
	<synonym relationship='synonym'>Gasterosteus neustrianus Blanchard, 1866</synonym>
	<synonym relationship='synonym'>Gasterosteus niger Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus noveboracensis Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus obolarius Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus plebeius Girard, 1854</synonym>
	<synonym relationship='synonym'>Gasterosteus ponticus Nordmann, 1840</synonym>
	<synonym relationship='synonym'>Gasterosteus pugetti Girard, 1856</synonym>
	<synonym relationship='synonym'>Gasterosteus quadrispinosa Crespon, 1844</synonym>
	<synonym relationship='synonym'>Gasterosteus quadrispinosus Crespon, 1844</synonym>
	<synonym relationship='synonym'>Gasterosteus santaeannae Regan, 1909</synonym>
	<synonym relationship='synonym'>Gasterosteus semiarmatus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus semiloricatus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus serratus Ayres, 1855</synonym>
	<synonym relationship='synonym'>Gasterosteus spinulosus Yarrell, 1835</synonym>
	<synonym relationship='synonym'>Gasterosteus suppositus Sauvage, 1874</synonym>
	<synonym relationship='synonym'>Gasterosteus teraculeatus Lacepède, 1801</synonym>
	<synonym relationship='synonym'>Gasterosteus tetracanthus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus texanus Sauvage, 1874</synonym>
	<synonym relationship='synonym'>Gasterosteus trachurus Cuvier, 1829</synonym>
	<synonym relationship='synonym'>Gasterosteus williamsoni Girard, 1854</synonym>
	<synonym relationship='synonym'>Gastrosteus hologymnus Regan, 1909</synonym>
	<synonym relationship='synonym'>Leiurus aculeatus (Linnaeus, 1758)</synonym>
	<dcterms:created>1991-03-05</dcterms:created>
	<dcterms:modified>2013-09-19</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118023&amp;speccode=2420'>Bakhvalova, A.E., T.S. Ivanova, M.V. Ivanov, A.S. Demchuk, E.A. Movchan and D.L. Lajus0 Long-term changes in the role of threespine stickleback (&lt;i&gt;Gasterosteus aculeatus&lt;/i&gt;) in the White Sea: predatory fish consumption reflects fluctuating stickleback abundance during the last century. Evolutionary Ecology Research 17(3):317-334. (Ref. 118023)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=2420'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='978-1-55362' url='http://www.fishbase.org/references/FBRefSummary.php?id=119011&amp;speccode=2420'>GoY, Government of Yukon0 Yukon freshwater fishes. Wildlife Viewing Program, Environment Yukon. Whitehorse (YU, Canada): Government of Yukon, 32 p. (Ref. 119011)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119049&amp;speccode=2420'>Antunes, C., F. Cobo and M.J. Araujo0 Iberian inland fisheries. p. 268-282. In J.F. Craig (ed.).  Freshwater Fisheries Ecology. John Wiley &amp; Sons Ltd. 899p. (Ref. 119049)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119063&amp;speccode=2420'>Norin, T. and T.D. Clark0 Measurement and relevance of maximum metabolic rate in fishes. J. Fish Biol. 88(1):122-151. (Ref. 119063)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119194&amp;speccode=2420'>Tonn, W., H. Swanson, C. Paszkowski, J. Hanisch and L. Chavarie0 Northern North America. p. 85-100. In J.F. Craig (ed.).  Freshwater Fisheries Ecology. John Wiley &amp; Sons Ltd. 899p. (Ref. 119194)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119390&amp;speccode=2420'>Yershov, P. and A. Sukhotin0 Age and growth of marine three-spined stickleback in the White Sea 50 years after a population collapse. Polar Biology 38(11):1813-1823. (Ref. 119390)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=120407&amp;speccode=2420'>Wienerroither R., Johannesen E., Dolgov A., Byrkjedal I., Aglen A., Bjelland O., Drevetnyak K., Eriksen KB., Høines Å., Langhelle G., Langøy H., Murashko P., Prokhorova T., Prozorkevich D., Smirnov O., Wenneck T.0 Atlas of the Barents Sea Fishes based on the winter survey. IMR-PINRO Joint Report Series 2-2013:220 pp. (Ref. 120407)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121185&amp;speccode=2420'>Dronen, N.O., C.K. Blend, M.C. Ostrowski de Núñez, S.K. Malhotra, N. Jaiswal0 Keys to the species of &lt;i&gt;Neolebouria&lt;/i&gt; Gibson, 1976 (Digenea: Opecoelidae: Plagioporinae), with a redescription of &lt;i&gt;Neolebouria truncata&lt;/i&gt; (Linton, 1940). Systematic Parasitology 88(3):213-225. (Ref. 121185)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121227&amp;speccode=2420'>Regan, C.T.0 A revision of the clupeid fishes of the genera &lt;i&gt;Sardinella&lt;/i&gt;, &lt;i&gt;Harengula&lt;/i&gt;, &amp; c. Ann. Mag. Nat. Hist. (Ser. 8) 19:377-395. (Ref. 121227)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121277&amp;speccode=2420'>Thorsteinson, L.K. and M.S. Love0 Alaska Arctic marine fish ecology catalog. Scientific Investigations Report 2016-5038. (Ref. 121277)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122793&amp;speccode=2420'>Tammiksaar, E. and K. Kangur0 Fish and fishing in Lake Peipsi (Estonia/Russia) since 1851: Similarities and differences between historical and modern times. J. Great Lakes Res. 46(2020):862-869. (Ref. 122793)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123027&amp;speccode=2420'>Bolshakova, Y.Y. and D.V. Bolshakov0 Ichthyofauna of the eastern coast bays of the Novaya Zemlya Archipelago. Oceanology 58(2):228-232. (Ref. 123027)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124235&amp;speccode=2420'>Ivanova, T.S., M.V. Ivanov, P.V. Golovin, N.V. Polyakova and D.L. Lajus0 The White Sea threespine stickleback population: Spawning habitats, mortality, and abundance. Evolutionary Ecology Research 17:301-315. (Ref. 124235)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=125501&amp;speccode=2420'>Dhora, D.0 Updated lists of the fish species of Albania's freshwater. Bul. Shk. Ser. Shk. Nat. 2020(70):46-74. (Ref. 125501)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=2420'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=2420'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132183&amp;speccode=2420'>Simian, G., D. Abraham, N. Bailly, P. Béarez, F. Urtizberea, A. Perrot, H. Goraguer, R. Langlois, P. Daszkiewicz, F. Teletchea, S. Tercerie and G. Denys0 Checklist of fishes from the Saint-Pierre and Miquelon archipelago. Cybium 46(4):371-384. (Ref. 132183)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132215&amp;speccode=2420'>WDNR, Wisconsin Department of Natural Resources0 Wisconsin Fish Species List.. World Wide Web electronic publication, Accessed 05 Oct. 2024. Madison (USA, WI):  Wisconsin Department of Natural Resources, 6 p. https://dnr.wisconsin.gov/sites/default/files/topic/Fishing/Species_wifish.pdf (Ref. 132215)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=2420'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=2420'>Corush, J.B.0 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference><dataObject>
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	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2420</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle</subject>
	<dc:description>Adults occur in fresh waters, estuaries and coastal seas (Ref. 4119).  Anadromous, with numerous non-anadromous populations in brackish or pure freshwater, rarely in marine waters.  In the sea, confined to coastal waters.  In freshwater, adults prefer to live in small stream but may occur in a variety of habitats including lakes and large rivers (Ref. 59043).  Inhabit shallow vegetated areas, usually over mud or sand (Ref. 5723).  Form schools.  Young associated with drifting seaweed (Ref. 12114, 12115).  Juveniles move to the sea (anadromous populations) or to deeper, larger water bodies (freshwater populations) in July-August, forming large feeding schools (Ref. 59043).  Feed on worms, crustaceans, larvae and adult aquatic insects, drowned aerial insects, and small fishes; has also been reported to feed on their own fry and eggs (Ref. 1998).  Eggs are found in nests constructed from plant material (Ref. 41678).  Males build, guard and aerate the nest where the eggs are deposited (Ref. 205).  Maximum length in freshwater is 8 cm while in saltwater is 11 cm (Ref. 35388).  Occasionally taken commercially in Scandinavia and processed into fishmeal and oil (Ref. 28219, 28964).  Commonly used as a laboratory animal (Ref. 1998).  A large bibliography is available at www.geocities.com/CapeCanaveral/Hall/1345/stickbibl.html.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86798&amp;speccode=2420'>Page, L.M. and B.M. Burr 2011 A field guide to freshwater fishes of North America north of Mexico. Boston : Houghton Mifflin Harcourt, 663p. (Ref. 86798)</reference></dataObject><dataObject>
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	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
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	<dc:description>Distinguished uniquely from its congeners in Europe by having trunk and caudal peduncle covered by a complete series of 29-35 bony scutes.  Other characters important to separate this species from other species of the genus include posterior edge of scutes crenulated and scutes forming a lateral keel on caudal peduncle.  Scutes may be missing on posterior part of trunk in hybrid zone with Gasterosteus gymnurus and in some isolated freshwater populations of northeastern Europe (Ref. 59043).  Identified by the 3 to 4 sharp, free spines before the dorsal fin, the pelvic fin reduced to a sharp spine and a small ray, and the series of plates along the sides of the body (Ref. 27547).  Gill rakers long and slender, 17 to 25 on the first arch or strictly freshwater forms, 1 or 2 more in anadromous forms; lateral line with microscopic pores (Ref. 27547).  The anadromous form is fully plated, with up to 37 plates on the sides and a rather pronounced keel on each side of the caudal peduncle (Ref. 27547).  Dorsal spines separated from each other and from the soft-rayed fins, each spine having a reduced membrane attached to its posterior side; anal spine free from rest of the fin; posterior margin of pectorals nearly truncate; caudal truncate to slightly indented (Ref. 27547).  Freshwater forms usually mottled brown or greenish; anadromous forms silvery green to bluish black (Ref. 27547).  A few isolated populations are black (Ref. 27547).  Sides usually pale; belly yellow, white or silvery (Ref. 27547).  Fins pale; pectoral rays often have dark dots (Ref. 27547).  Breeding males (except for black forms) become brilliant bluish or green with blue or green eyes, and the forward part of the body, especially the breast region, turns bright red or orange (Ref. 27547).  Caudal fin with 12 rays (Ref. 2196).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4119&amp;speccode=2420'>Banister, K. 1986 Gasterosteidae. p. 640-643. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the north-eastern Atlantic and the Mediterranean. volume 2. UNESCO, Paris. (Ref. 4119)</reference></dataObject><dataObject>
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	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Dorsal spines (total): 2 - 4; Dorsal soft rays (total): 10 - 14;  Analspines: 1; Analsoft rays: 7 - 11; Vertebrae: 29 - 34</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4119&amp;speccode=2420'>Banister, K. 1986 Gasterosteidae. p. 640-643. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the north-eastern Atlantic and the Mediterranean. volume 2. UNESCO, Paris. (Ref. 4119)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Circumarctic and temperate regions:  Extending south to the Black Sea, southern Italy, Iberian Peninsula, North Africa; in Eastern Asia north of Japan (35°N), in North America north of 30-32°N; Greenland.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12841&amp;speccode=2420'>Monod, T. 1979 Gasterosteidae. p. 280-286. In J. C. Hureau and Th. Monod (eds.) Check-list of the Fishes of the north-eastern Atlantic and of the Mediterranean (CLOFNAM). UNESCO, Paris. Vol. 1. (Ref. 12841)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
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	<dc:title>Importance</dc:title>
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	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2420</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>fisheries: minor commercial; aquarium: public aquariums; price category: unknown; price reliability: </dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27547&amp;speccode=2420'>Morrow, J.E. 1980 The freshwater fishes of Alaska. University of. B.C. Animal Resources Ecology Library. 248p. (Ref. 27547)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1672&amp;speccode=2420'>Riehl, R. and H.A. Baensch 1991 Aquarien Atlas. Band. 1. Melle: Mergus, Verlag für Natur-und Heimtierkunde, Germany. 992 p. (Ref. 1672)</reference></dataObject><dataObject>
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	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2420</dc:source>
	<dc:description>Anadromous. Fish that ascend rivers to spawn, as salmon and hilsa do. Sub-division of diadromous.    Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=2420'>Corush, J.B. 2019 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=2420&amp;StockCode=2615</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Spawning behavior is similar for both freshwater and anadromous forms (Ref. 28966). Just before breeding, males become very territorial. The male builds a nest of plant-material glued together with spiggin, a protein produced in the kidney (Ref. 52349). Once a nest is built, the male entices the female into the nest by performing a courtship dance which is a series of  zigzag movements (Ref. 1998). A receptive female follows the male who points the opening of the nest by posing above it with his head down. The female enters the nest, deposits up to a few hundred eggs, and is driven out by the male after eggs have been deposited. The male then enters the nest to fertilize the eggs. The male can choose to court another female to enter the nest and lay eggs before entering himself to fertilise the deposited eggs. Females may lay eggs in several nests over a period of several days or may be courted by the same male (Ref. 27547). The male guards and ventilates the eggs and young (Ref. 1998).  During spawning season, males develop a bright orange to red belly and blue-green flank and eyes.  Eggs hatch in 7-8 days.  Anadromous forms usually die of exhaustion after spawning cycle.  Freshwater individuals are able to complete several cycles within one year or sometimes over several years (Ref. 59043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7471'>Balon, E.K. 1990 Epigenesis of an epigeneticist: the development of some alternative concepts on the early ontogeny and evolution of fishes. Guelph Ichthyol. Rev. 1:1-48.</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-2420</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
	<dcterms:modified>2007-08-01 00:00:00</dcterms:modified>
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	<dc:source>aculeatus</dc:source>
	<dc:description>Shoaling species outside the breeding season, especially when young.  Inhabits vegetated areas, usually over mud or sand (Ref. 5723).  In the sea, confined to coastal waters.  Forms schools.  Eggs are found in nests constructed from plant material (Ref. 41678).  Anadromous and nerito-pelagic (Ref. 58426).  Feeds on worms, crustaceans, larvae and adult aquatic insects, drowned aerial insects, and small fishes; has also been reported to feed on their own fry and eggs (Ref. 1998).  Length in freshwater is 8 cm while in saltwater is 11 cm (Ref. 35388).  Preyed upon by American mergansers.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4119&amp;speccode=2420'>Banister, K. 1986 Gasterosteidae. p. 640-643. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the north-eastern Atlantic and the Mediterranean. volume 2. UNESCO, Paris. (Ref. 4119)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2420</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle</subject>
	<dc:description>11.0 cm TL (male/unsexed; (Ref. 35388)); max. reported age: 8 years (Ref. 72489)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='87-90787-00-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=35388&amp;speccode=2420'>Muus, B.J. and J.G. Nielsen 1999 Sea fish. Scandinavian Fishing Year Book, Hedehusene, Denmark. 340 p. (Ref. 35388)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=72489&amp;speccode=2420'>Reimchen, T.E. 1992 Extended longevity in a large-bodied stickleback, &lt;i&gt;Gasterosteus&lt;/i&gt;, population. Canadian Field Naturalist 106:129-131. (Ref. 72489)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-2420</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-03-05 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-21 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2420</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>benthopelagic; anadromous (Ref. 138280); freshwater; brackish; marine; depth range 0 - 100 m (Ref. 50550)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138280&amp;speccode=2420'>Corush, J.B. 2019 Evolutionary patterns of diadromy in fishes: more than a transitional state between marine and freshwater. BMC Evol. Biol. 19(168):13 p. (Ref. 138280)</reference>
	<reference isbn='5-8044-0308-7' url='http://www.fishbase.org/references/FBRefSummary.php?id=50550&amp;speccode=2420'>Fedorov, V.V., I.A. Chereshnev, M.V. Nazarkin, A.V. Shestakov and V.V. Volobuev 2003 Catalog of marine and freswater fishes of the northern part of the Sea of Okhotsk. Vladivostok: Dalnauka, 2003. 204 p. (Ref. 50550)</reference></dataObject>
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