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	<dc:identifier>FB-7331</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Serranoidei</dwc:Order>
	<dwc:Family>Epinephelidae</dwc:Family>
	<dwc:Genus>Epinephelus</dwc:Genus>
	<dwc:ScientificName>Epinephelus bilobatus Randall &amp; Allen, 1987</dwc:ScientificName>
	<commonName xml:lang='en'>Brown-spotted rock-cod</commonName>
	<commonName xml:lang='en'>Frostback cod</commonName>
	<commonName xml:lang='en'>Frostback rockcod</commonName>
	<commonName xml:lang='en'>Twinspot grouper</commonName>
	<commonName xml:lang='fr'>Mérou géminé</commonName>
	<commonName xml:lang='he'>Lokos</commonName>
	<commonName xml:lang='es'>Mero de pintas gemelas</commonName>
	<commonName xml:lang=''>白背石斑魚</commonName>
	<commonName xml:lang=''>白背石斑鱼</commonName>
	<dcterms:created>1993-02-12</dcterms:created>
	<dcterms:modified>2017-06-06</dcterms:modified>
	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=7331'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1830&amp;speccode=7331'>Eschmeyer, W.N.0 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p. (Ref. 1830)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4787&amp;speccode=7331'>Randall, J.E. and P.C. Heemstra0 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall0 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=7331'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31667&amp;speccode=7331'>Anonymous0 Fish collection database of the Bernice P. Bishop Museum (BPBM). Bishop Museum, 1525 Bernice Street, Honolulu, Hawai`I, 96817-0916 USA. (Ref. 31667)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39231&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall0 Serranidae. p. 2442-2547. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 4. Bony fishes part 2 (Mugilidae to Carangidae). Rome, FAO. 2069-2790 p. (Ref. 39231)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=7331'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47567&amp;speccode=7331'>Allen, G.R. and M. Adrim0 Coral reef fishes of Indonesia. Zool. Stud. 42(1):1-72. (Ref. 47567)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=7331'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58534&amp;speccode=7331'>Nguyen, N.T. and V.Q. Nguyen0 Biodiversity and living resources of the coral reef fishes in Vietnam marine waters. Science and Technology Publishing House, Hanoi. (Ref. 58534)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75154&amp;speccode=7331'>Hoese, D.F., D.J. Bray, J.R. Paxton and G.R. Allen0 Fishes. In Beasley, O.L. and A. Wells (eds.) Zoological Catalogue of Australia. Volume 35 Australia: ABRS &amp; CSIRO Publishing, 2178 p. (Ref. 75154)</reference>
	<reference isbn='9781466506022' url='http://www.fishbase.org/references/FBRefSummary.php?id=89707&amp;speccode=7331'>Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra0 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707)</reference>
	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=7331'>Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115074&amp;speccode=7331'>Anderson, E.S.0 The response of a predatory fish, &lt;i&gt;Ophiodon elongatus&lt;/i&gt;, to a marine protected area: variation in diet, catch rates, and size composition. California Polytechnic State University. (Ref. 115074)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115274&amp;speccode=7331'>Clough, J.M.0 Demersal fish assemblages of South Passage and Blind Strait, Western Australia: a unique subtropical embayment in a World Heritage Property - RAW DATA in excel file.. University of Western Australia, 6 worksheets (Ref. 115274)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=119023&amp;speccode=7331'>Smallwood, C.B., A. Tate and K.L. Ryan0 Weight-length summaries for Western Australian fish species derived from surveys of recreational fishers at boat ramps. Fisheries Research Report No. 278, Department of Primary Industries and Regional Development, Western Australia. 151pp. (Ref. 119023)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122752&amp;speccode=7331'>University of Western Australia0 Benthic  BRUVS data 2019. Video BRUVS from Jessica Meeuwig. (Ref. 122752)</reference><dataObject>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=823' logoURL='' role='photographer'> FAO</agent>
	<dcterms:created>1996-05-25</dcterms:created>
	<dcterms:modified>2010-07-09</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights>This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged</dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7331</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description></dc:description>
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	<dc:identifier>FB-GeneralDescr-7331</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1993-02-12 00:00:00</dcterms:created>
	<dcterms:modified>2017-06-06 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7331</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>No fishery information is available.  Found on coral reefs or rocky bottom (Ref. 089707).  No published information on its biology. Solitary (Ref 90102).</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-7331</dc:identifier>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
	<dcterms:created>1994-06-10 00:00:00</dcterms:created>
	<dcterms:modified>2013-09-02 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7331</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>Characterized by:  greyish-brown color overall with hexagonal brown spots; white strip at back; dorsal fin base with 2-3 dark bilobed spots; body scales ctenoid, cycloid scales anterodorsally on thorax and ventrally on abdomen; body with auxiliary scales; greatest depth of body 2.8-3.3 in SL; rounded caudal fin; pelvic fins 1.8-2.1 in head length (Ref. 90102); preopercle subangular, with serrae at the angle slightly enlarged; opercular spines inconspicuous; straight or slightly concave upper edge of operculum; nostrils subequal in small fish, but posterior nostril diameter about twice of that of anterior nostril in a fish of 27 cm SL; maxilla reaches below rear half of eye; 2 rows of small teeth on midlateral part of lower jaw (Ref. 089707).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-7331</dc:identifier>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
	<dcterms:created>1994-06-10 00:00:00</dcterms:created>
	<dcterms:modified>2013-09-02 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7331</dc:source>
	<dc:description>Dorsal spines (total): 11; Dorsal soft rays (total): 17 - 18;  Analspines: 3; Analsoft rays: 8</dc:description>
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	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-7331</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1993-02-12 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Eastern Indian Ocean:  known only from Western Australia.  Was illustrated in color as Epinephelus maculatus by Allen (1985, Ref. 1677) and Sainsbury et al.  (1985, Ref. 3131).  This species have been much confused with other 'reticulated groupers' in literature, and many museum specimens have been misidentified.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-7331</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Auda Kareen Ortañez</agent>
	<dcterms:created>2007-08-08 00:00:00</dcterms:created>
	<dcterms:modified>2008-11-18 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>bilobatus</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses</subject>
	<dc:description>Occurs in inshore waters of the continental shelf (Ref. 75154).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58534&amp;speccode=7331'>Nguyen, N.T. and V.Q. Nguyen 2006 Biodiversity and living resources of the coral reef fishes in Vietnam marine waters. Science and Technology Publishing House, Hanoi. (Ref. 58534)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-7331</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1993-02-12 00:00:00</dcterms:created>
	<dcterms:modified>2017-06-06 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7331</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>33.0 cm TL (male/unsexed; (Ref. 5222))</dc:description>
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	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-7331</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1993-02-12 00:00:00</dcterms:created>
	<dcterms:modified>2017-06-06 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7331</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>reef-associated; marine; depth range 4 - 50 m (Ref. 5222)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=7331'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject>
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