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	<dc:identifier>FB-8932</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8932</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Coptodon</dwc:Genus>
	<dwc:ScientificName>Coptodon tholloni (Sauvage, 1884)</dwc:ScientificName>
	<commonName xml:lang=''>Ekouni</commonName>
	<commonName xml:lang='fi'>Kirjotilapia</commonName>
	<commonName xml:lang='fr'>Carpe</commonName>
	<commonName xml:lang=''>Ompoumini</commonName>
	<commonName xml:lang=''>索氏非鯽</commonName>
	<commonName xml:lang=''>索氏非鲫</commonName>
	<synonym relationship='synonym'>Chromis ogowensis Günther, 1896</synonym>
	<synonym relationship='synonym'>Chromis tholloni Sauvage, 1884</synonym>
	<synonym relationship='misapplied name'>Tilapia melanopleura (non Duméril, 1861)</synonym>
	<synonym relationship='synonym'>Tilapia ogowensis (Günther, 1896)</synonym>
	<synonym relationship='synonym'>Tilapia tholloni (Sauvage, 1884)</synonym>
	<synonym relationship='synonym'>Tilapia tholloni tholloni (Sauvage, 1884)</synonym>
	<dcterms:created>1994-03-04</dcterms:created>
	<dcterms:modified>2024-03-22</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=8932'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8932'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12468&amp;speccode=8932'>Baensch, H.A. and R. Riehl0 Aquarien Atlas. Band 4. Mergus Verlag GmbH, Verlag für Natur-und Heimtierkunde, Melle, Germany. 864 p. (Ref. 12468)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=8932'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=8932'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8932'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=8932'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=8932'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50787&amp;speccode=8932'>Ogandagas, C.0 National Report. Marine biodiversity in Gabon - the known and the unknown. p. 75-85. In C. Decker, C. Griffiths, K. Prochazka, C. Ras and A. Whitfield (eds.) Marine biodiversity in sub-saharan Africa: the known and unknown. (Ref. 50787)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=8932'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='2.11.086730-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=51193&amp;speccode=8932'>Vivien, J.0 Faune du Cameroun. Guide des mammifères et poissons. GICAM et Ministère de la Coopération et du Développement, Yaoundé/Paris. 271 p. (Ref. 51193)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=8932'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52869&amp;speccode=8932'>Mve, R.O. and J.D. Mbega0 Noms communs d'espèces importantes de poisson en langue Fang, Gabon. p. 243-245. In M.L.D. Palomares, B. Samb, T. Diouf, J.M. Vakily and D. Pauly (eds.) Fish biodiversity: local studies as basis for global inferences. ACP-EU Fish. Res. Rep. 14:281 p. (Ref. 52869)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55074&amp;speccode=8932'>Thys van den Audenaerde, D.F.E.0 Révision systématique des espèces congolaises du genre &lt;i&gt;Tilapia&lt;/i&gt; (Pisces, Cichlidae). Ann. Mus. R. Afr. Centr., Sér. In-8°, Sci. Zool., 124:155p. (Ref. 55074)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8932'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=74387&amp;speccode=8932'>Thys van den Audenaerde, D.F.E.0 La distribution géographique des &lt;i&gt;Tilapia&lt;/i&gt; au Congo. Bull. Séances Acad. R. Sci. Outre-Mer 9(3):570-605. (Ref. 74387)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=8932'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels0 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81651&amp;speccode=8932'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels0 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G. Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Volume II. Collection Faune et Flore tropicales 42. Institut de Recherche pour le Développement, Paris, France, Muséum National d’Histoire Naturelle, Paris, France, and Musée Royal de l’Afrique Centrale, Tervuren, Belgium. 603 pp. (Ref. 81651)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=8932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=8932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=8932'>Dunz, A.R. and U.K. Schliewen0 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93897&amp;speccode=8932'>Monsembula Iyaba, R.J.C. and M.L.J. Stiassny0 Fishes of the Salonga National Park (Congo basin, central Africa): a list of species collected in the Luilaka, Salonga, and Yenge Rivers (Equateur Province, Democratic Republic of Congo). Check List 9(2):246-256. (Ref. 93897)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=8932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=8932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='978-3-030-03083-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=120641&amp;speccode=8932'>Skelton, P.H.0 Chapter 11. The freshwater fishes of Angola. p. 207-242. In B.J. Huntley, V. Russo, F. Lages and N. Ferrand (eds.) Biodiversity of Angola. Science &amp; conservation: a modern synthesis. Springer Nature Switzerland AG, Cham, Switzerland. 549 p. (Ref. 120641)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=8932'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference><dataObject>
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	<dc:description>Pair-bonding, open (Ref. 52307) substrate brooder (Ref. 52307, 81260) with both parents guarding the brood (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=8932'>Dunz, A.R. and U.K. Schliewen 2013 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference></dataObject><dataObject>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
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	<dc:description>Diagnosis: 10 or fewer rakers on lower limb of first arch; unpaired fins not predominantly red-brown; mouth terminal; caudal fin densely covered with scales and with network of clear spots and lines; at least 1 row (sometimes 2) of dark spots in the middle of the body; a black sickle-shaped spot at the base of each flank scale; dorsal fin with 15-16 spines and 9-11 soft rays (total rays 24-26); anal fin with 3 spines and 7-9 soft rays; 27-29 scales in lateral line; outer row teeth relatively fine (Ref. 81260).Description: relatively slender-bodied; mouth large; head profile of adults, especially large males, slightly concave; eyes relatively large (Ref. 52307). Head length 32.3-35.0% SL; lower pharyngeal jaw with pointed and curved anterior teeth and straight tricuspid posterior teeth; ventral keel shorter than dentigerous plate; 2-4 tooth rows in oral jaws, outer row teeth relatively fine, bicuspids (Ref. 81260).Coloration: flanks yellowish white, creamy white ventrally, dark brown dorsally; flank scales with a black marking at the base; belly, throat and chest whitish in juveniles, becoming red in sexually mature specimens; 7 vertical bars, narrowing ventrally, transversed by 2 horizontal bands; where bars and bands intersect they form a series of blotches; dorsal fin greyish with some clear maculae and a well marked "tilapia spot"; caudal fin dark with numerous maculae, particularly near the distal border; anal fin very dark to black, sometimes tinted with red; pelvics with black leading edge; pectoral fins transparent (Ref. 81260). A black band crosses through the eye (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=8932'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels 2008 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 15 - 16; Dorsal soft rays (total): 9 - 11;  Analspines: 3; Analsoft rays: 7 - 9</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81260&amp;speccode=8932'>Stiassny, M.L.J., A. Lamboj, D. De Weirdt and G.G. Teugels 2008 Cichlidae. p. 269-403. In M.L.J. Stiassny, G.G Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa Volume 2. Coll. faune et flore tropicales 42. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 603p. (Ref. 81260)</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Africa:  from the Ogooué River in Gabon (Ref. 81260) to the lower Congo River in Democratic Republic of the Congo (Ref. 55074, 74387, 120641), including the Nyanza in Gabon, Kouilou-Niari in Republic of Congo, and the Chiloango in Angola and Democratic Republic of the Congo (Ref. 81260). Specimens from Pool Malebo are either this species or C. congica (Ref. 55074, 74387).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8932'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:description>22.0 cm TL (male/unsexed; (Ref. 5163))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8932'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:description>benthopelagic; freshwater</dc:description>
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